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Overview Fotosintesis
Dr. Riris Istighfari Jenie, M.Si., Apt.
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Energi dapat diubah dari satu
bentuk ke bentuk lainnyaEnergi bebas
(available for work)
vs.
Panas(not available for work)
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Matahari: Sumber energi utama
kehidupan di bumi
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• Hampir semua tanaman adalah autotrof fotosintetik,
seperti halnya beberapa bakteria dan protists. – Autotrof menghasilkan senyawa-senyawa organiknya
sendiri melalui fotosintesis.
– Energi cahaya matahari diubah menjadi energi yang
tersimpan dalam bentuk ikatan kimia.
(a) Mosses, ferns, andf lowering plants
(b) Kelp
(c) Euglena (d) Cyanobacteria
Dasar-dasar Fotosintesis
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Light Energy Harvested by Plants &
Other Photosynthetic Autotrophs
6 CO2 + 6 H2O + light energy → C6H12O6 + 6 O2
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Food Chain
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THE FOOD WEB
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WHY ARE PLANTS GREEN?
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Electromagnetic Spectrum and
Visible LightGamma
rays X-rays UVInfrared &
Microwaves Radio waves
Visible light
Wavelength (nm)
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Different wavelengths of visible light are seen bythe human eye as different colors.
WHY ARE PLANTS GREEN?
Gamma
raysX-rays UV Infrared
Micro-
waves
Radio
waves
Visible light
Wavelength (nm)
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Sunlight minus absorbedwavelengths or colors
equals the apparent color
of an object.
The feathers of male cardinals
are loaded with carotenoid
pigments. These pigments
absorb some wavelengths of
light and reflect others.
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Why are plants green?
Transmitted light
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WHY ARE PLANTS GREEN?
Plant Cellshave Green
Chloroplasts
The thylakoid
membrane of the
chloroplast isimpregnated with
photosynthetic
pigments (i.e.,
chlorophylls,
carotenoids).
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• Chloroplasts
absorb light
energy andconvert it to
chemical energy
LightReflected
light
Absorbedlight
Transmittedlight
Chloroplast
THE COLOR OF LIGHT SEEN IS THE
COLOR NOT ABSORBED
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• Fotosintesis adalah proses dimana
organisme autotrof menggunakan energi
sinar matahari untuk menghasilkan gula dan
oksigen dari karbon dioksida dan air.
Overview Fotosintesis
Carbondioxide
Water Glucose Oxygengas
PHOTOSYNTHESIS
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• The Calvin cycle makessugar from carbon
dioxide – ATP generated by the light
reactions provides the energyfor sugar synthesis
– The NADPH produced by the
light reactions provides the
electrons for the reduction of
carbon dioxide to glucose
Light
Chloroplast
Lightreactions
Calvincycle
NADP
ADP+ P
• The light reactions
convert solarenergy to chemicalenergy – Produce ATP & NADPH
Overview Fotosintesis
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Chloroplasts: Sites of Photosynthesis
• Photosynthesis – Occurs in chloroplasts, organelles in certain
plants
– All green plant parts have chloroplasts and carryout photosynthesis
• The leaves have the most chloroplasts
• The green color comes from chlorophyll in the
chloroplasts• The pigments absorb light energy
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• In most plants, photosynthesis occursprimarily in the leaves, in the chloroplasts
• A chloroplast contains:
– stroma, a fluid – grana, stacks of thylakoids
• The thylakoids contain chlorophyll
– Chlorophyll is the green pigment that captureslight for photosynthesis
Photosynthesis occurs in chloroplasts
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• The location and structure of chloroplasts
LEAF CROSS SECTIONMESOPHYLL CELL
LEAF
Chloroplast
Mesophyll
CHLOROPLAST Intermembrane space
Outer membrane
Inner membrane
ThylakoidcompartmentThylakoidStroma
Granum
StromaGrana
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• Chloroplasts contain several pigments
Chloroplast Pigments
– Chlorophyll a
– Chlorophyll b
– Carotenoids
Figure 7.7
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Chlorophyll a & b•Chl a has a methyl
group
•Chl b has a carbonyl
group
Porphyrin ring
delocalized e-
Phytol tail
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Different pigments absorb light
differently
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Excitedstate
e
Heat
Light
Photon
Light(fluorescence)
Chlorophyll
molecule
Groundstate
2
(a) Absorption of a photon
(b) fluorescence of isolated chlorophyll in solution
Eksitasi klorofil di dalam
sebuah kloroplas
Hilangnya energi karena panas
menyebabkan foton cahaya
menjadi berenergi lebih rendah.
Energi yang lebih rendah berarti
lebih panjang gelombangcahayanya.
Energy = (Planck’s constant) x
(velocity of light)/(wavelength of
light)
Bergeser ke cahaya warnamerah di spektrum cahaya.
e
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Cyclic Photophosphorylation• Process for ATP generation associated with
some Photosynthetic Bacteria• Reaction Center => 700 nm
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Primaryelectron acceptor
Primaryelectron acceptor
Photons
PHOTOSYSTEM I
PHOTOSYSTEM II
Energy for synthesis of
by chemiosmosis
Noncyclic Photophosphorylation• Photosystem II regains electrons by splitting
water, leaving O2 gas as a by-product
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• The O2 liberated by photosynthesis is madefrom the oxygen in water (H+ and e-)
Plants produce O2 gas by splitting H2O
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2 H + 1 /2
Water-splittingphotosystem
Reaction-center
chlorophyll
Light
Primaryelectronacceptor
Energyto make
Primaryelectronacceptor
Primaryelectronacceptor
NADPH-producingphotosystem
Light
NADP
1
2
3
How the Light Reactions Generate ATP and NADPH
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Water-splittingphotosystem
NADPH-producingphotosystem
ATPmill
• Two types of
photosystems
cooperate in the
light reactions
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• Dua fotosistem yang berhubungan
mengumpulkan foton cahaya matahari dan
mentransfer energi tsb ke elektron klorofil.
• Elektron-elektron tereksitasi diteruskan dari
akseptor elektron primer ke rantai transpor
elektron. – Pada akhirnya akan diperoleh ATP dan NADPH
Pada reaksi cahaya, transpor elektron
menghasilkan ATP, NADPH, & O2
K i i kk i t i
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• Rantai transpor elektron diatur melalui
fotosistem di membran tilakoid dan
memompa H+ melewati membran tersebut.
– Aliran balik H+ melalui membran dimanfaatkan
oleh ATP sintase untuk membentuk ATP
– Di stroma, ion H+ bergabung NADP+ membentuk
NADPH
Kemiosmosis menggerakkan sintesis
ATP pada reaksi cahaya
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• Produksi ATP oleh kemiosmosis dalam
fotosintesis
Thylakoidcompartment(high H+)
Thylakoidmembrane
Stroma(low H+)
Light
Antennamolecules
Light
ELECTRON TRANSPORT
CHAIN
PHOTOSYSTEM II PHOTOSYSTEM I ATP SYNTHASE
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• A Photosynthesis Road Map
Chloroplast
Light
Stack of thylakoids ADP
+ P
NADP
Stroma
Lightreactions
Calvincycle
Sugar used for
Cellular respiration
Cellulose
Starch
Other organic compounds
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Review: Photosynthesis uses light
energy to make food molecules
Light
Chloroplast
Photosystem IIElectrontransport
chainsPhotosystem I
CALVINCYCLE Stroma
LIGHT REACTIONS CALVIN CYCLE
Cellularrespiration
Cellulose
Starch
Otherorganiccompounds
• A summary of
the chemical
processes ofphotosynthesis
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It's not that
easy bein'
green… but itis essential for
life on earth!